Middle Pleistocene cold stage climates in the Mediterranean: New evidence from the glacial record

A major reduction in summer temperatures during a Middle Pleistocene glacial cycle caused the most extensive glaciation recorded in the Mediterranean region. Glaciers in the mountains of Greece formed during marine isotope stage (MIS) 12 (474 000–427 000 years BP) under climatic conditions characterised by mean summer temperatures at least 11 °C cooler than today and annual precipitation of ≤ 2300 mm at the equilibrium line altitude (1741 m a.s.l.). This represents the coldest mean summer temperatures recorded in Greece during at least the last 430 000 years. Later Pleistocene glaciations (MIS 6 and 5d-2) were characterised by warmer summer temperatures and higher annual precipitation.