Artinskian–Wordian antitropical rugose coral associations: A palaeogeographical approach

Recent investigations in the Urals and East-European Platform as well as study of corals collections from South Primorye (Russian Far East) and South-Eastern Pamirs have been undertaken. Antitropical rugose associations include taxa that are bounded in their distribution by the temperate zones of the Boreal and Perigondwanan realms. They occur sporadically in tropical latitudes but never form abundant and diverse communities. Antitropical associations are represented by the so-called ‘Cyathaxonia fauna’ (sensu lato), predominating in two selected stratigraphic intervals. The abrupt change of taxonomic composition from colonial to ‘Cyathaxonia fauna’ allows the separation of two assemblages: (i) mid Artinskian–Kungurian and (ii) Roadian–Wordian. The key regions for comparison are the Urals (mid Artinskian–Kungurian), Spitsbergen (Kungurian–Wordian), East-European Platform (Roadian), Texas (mid Artinskian–Kungurian), South Primorye (Wordian–Capitanian), South-Eastern Pamirs (Upper Artinskian), and the Sverdrup Basin (Wordian). The Late Artinskian fauna of the South-Eastern Pamirs and Wordian faunas of South Primorye bear features of mixed assemblages. A few models of the origination of antitropical assemblages are acceptable: (i) the convergent parallel evolution of Lophophyllidium in the Northern Hemisphere and Southern Hemisphere, (ii) a similar evolution trend developed in parallel in three geographically distant regions: Verbeekiella, Pseudowannerophyllum, and “Pseudoverbeekiella”, and (iii) immigration of some species within temperate zones caused by current circulation changes both in the Northern and Southern Hemispheres, as well as northward spreading of terranes in the Perigondwanan Realm.