Sympatry of two ankylosaurs (Hungarosaurus and cf. Struthiosaurus) in the Santonian of Hungary

A complete and well-preserved right ankylosaurian humerus from the Upper Cretaceous (Santonian) Csehbánya Formation of Iharkút, western Hungary is described here. Based on its osteological features and 21.5 cm adult length, the new specimen is markedly different from the slender humerus of Hungarosaurus, the previously known ankylosaur from the locality, and more similar to that of Struthiosaurus. Thus, the new Hungarian specimen is tentatively assigned here to cf. Struthiosaurus thereby dating back the first occurrence of this genus to the Santonian. The new fossil demonstrates the sympatric co-existence of two different nodosaurid ankylosaurs (a smaller, robust form with 2–2.5 m total body length and a larger, cursorial form with 4–4.5 m body length) in the Iharkút fauna. This also suggests that the pattern of the European ankylosaur diversity was more complex than previously thought.