A chirothere footprint from the Otter Sandstone Formation (Middle Triassic, late Anisian) of Devon, UK

The Middle Triassic (Anisian) Otter Sandstone Formation of Devon is well known as a source of vertebrate skeletal remains, particularly of reptiles. Here we report the first definite vertebrate trace fossil, a well-preserved chirothere footprint of uncertain taxonomic identity. Chirothere prints are mostly attributed to rauisuchian archosaurs, probable fragmentary remains of which have previously been recovered from the Otter Sandstone Formation.