Ten-year responses of ponderosa pine plantations to repeated vegetation and nutrient control along an environmental gradient

Factorial combinations of vegetation, nutrient, and insect control treatments were applied repeatedly to three contrasting California plantations of Pinus ponderosa var. ponderosa Dougl. ex Laws. Ten-year findings show that potential productivity is far greater than previously believed. Stem volume gains were linked directly with increases in crown volume. Insect problems were negligible. Vegetation control increased tree growth profoundly on xeric sites but less so on the most mesic. Where soil was both droughty and infertile, growth responses traced primarily to improved soil moisture availability and secondarily to better nutrition. The most fertile site also was droughty, and trees responded only to improved moisture availability. Water was less limiting on the most productive site. There, both fertilizers and herbicides triggered similar, substantive growth increases. Drought from both plant competition and climate reduced stomatal conductance, xylem water potential, and net assimilation rates. Assimilation rates increased linearly with site index, but treatment differences were not apparent once drought had peaked. Fertilization improved water-use efficiency where water stress was not extreme. Advantages in water availability to pines from vegetation control will dissipate as tree crowns close and transpiration rises.