The aftermath of tree death: coarse woody debris and the topography in four tropical rain forests

The relationship of coarse woody debris (CWD) was examined with respect to topography and site in four lowland tropical rain forests in northern Borneo and western Ecuador. In total, 1914 dead trees >20 cm diameter at breast height were enumerated in 46 ha. The basal area per hectare of dead trees varied strongly with both site and slope position. Dead basal area and its correlate dead tree density increased moving up the topographic gradient from the valleys to the ridges. Site estimates for CWD volume ranged from 96 to 154 m3·ha-1. Mean standing CWD volume was four times higher on the ridge tops (54 m3·ha-1) compared with the valley and cross-terrain areas (both 13 m3· ha-1). In contrast, downed volume did not vary with slope position. The proportions of CWD present as standing wood were two to three times higher in the Bornean sites compared with Hoja Blanca. These topographical and site differences in CWD were linked to differences in mode of tree death as well as the distribution of live trees per hectare.